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Class A. Leaves are evergreen, needlelike, narrow-linear, awl-shaped, or scalelike. |
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| Group ILeaves in bundles are needlelike. | |||||||||||||
| PinusPine | |||||||||||||
| Group IILeaves opposite and awl-shaped or scalelike. Leaves are small, often overlapping, and adpressed. | |||||||||||||
| I. Leaves are sharp and harsh to the touch, especially on the main branches or older twigs. Leaves are free at the apex. Fruit is berrylike. | |||||||||||||
| JuniperusJuniper | |||||||||||||
| II. Leaves are soft to the touch and scalelike. Most are closely adpressed to the branchlets. Fruit is a woody cone with six to 12 scales. | |||||||||||||
| ThujaArborvitae | |||||||||||||
| Group IIILeaves are alternate and scattered on the stem. Leaves are narrow and linear. | |||||||||||||
| I. Winter buds on normal shoots are conspicuously long-pointed, brown with many scales, and non-resinous. | |||||||||||||
| PseudotsugaDouglas-Fir | |||||||||||||
| II. Buds are not long-pointed. | |||||||||||||
| A. Leaves are deciduous at their base. Old twigs are not roughened by persistent leaf bases. | |||||||||||||
| AbiesFir | |||||||||||||
| AA. Leaves are deciduous above the leaf base, leaving the twigs roughened by persistent leaf bases. | |||||||||||||
| |. Leaves are flat and arranged into two planes. White, stomatic lines are present beneath the leaf. Twigs are only slightly roughened by persistent leaf bases. | |||||||||||||
| TsugaHemlock | |||||||||||||
| ||. Leaves are four-sided with stomatic lines of equal numbers on all sides. Twigs are strongly roughened by persistent leaf bases. | |||||||||||||
| PiceaSpruce | |||||||||||||
| Class B. Leaves are deciduous, mostly broad, and occasionally narrow or linear. | |||||||||||||
| Group ILeaves are opposite or whorled. | |||||||||||||
| Subgroup ILeaves are compound. | |||||||||||||
| I. Leaves are palmately compound. | |||||||||||||
| AesculusBuckeye, Horsechestnut | |||||||||||||
| II. Leaves are pinnately compound. | |||||||||||||
| A. Side buds are hidden. | |||||||||||||
| AcerBoxelder | |||||||||||||
| AA. Side buds are prominent. Bundle scars are numerous. | |||||||||||||
| FraxinusAsh | |||||||||||||
| Subgroup IILeaves are lobed. | |||||||||||||
| I. Pith is chambered or sometimes hollow. | |||||||||||||
| PaulowniaPaulownia | |||||||||||||
| II. Pith is solid. | |||||||||||||
| AcerMaple | |||||||||||||
| Subgroup IIILeaves are toothed. | |||||||||||||
| I. Petioles are connected directly or by a line around the twig. | |||||||||||||
| ViburnumViburnum | |||||||||||||
| II. Petioles are not connected either directly or by a line around the twig. | |||||||||||||
| EuonymusEuonymus | |||||||||||||
| Subgroup IVLeaves are entire. | |||||||||||||
| I. Pith is chambered or sometimes hollow. | |||||||||||||
| PaulowniaPaulownia | |||||||||||||
| II. Pith is solid. | |||||||||||||
| A. Leaves are long-petioled, large (412 inches [10301/2 cm] long), and often whorled. | |||||||||||||
| CatalpaCatalpa | |||||||||||||
| AA. Leaves are not as above. | |||||||||||||
| |. Bundle scars are almost in a complete circle. Side buds are globu- lar with four to five pairs of bud scales. | |||||||||||||
| ChionanthusFringetree | |||||||||||||
| ||. Bundle scars and buds are not as above. Leaves have parallel veins. | |||||||||||||
| CornusDogwood | |||||||||||||
| Group IILeaves are alternate. | |||||||||||||
| Subgroup ILeaves are compound. | |||||||||||||
| I. Leaves are twice compound. | |||||||||||||
| A. Twigs are spiny or thorny. | |||||||||||||
| |. Spines are unbranched, short, and stout. | |||||||||||||
| AraliaAralia | |||||||||||||
| ||. Thorn is simple or branched, stout, and 2-4 or more inches in length. | |||||||||||||
| GleditsiaHoneylocust | |||||||||||||
| AA. Twigs are not spiny or thorny. | |||||||||||||
| GymnocladusKentucky Coffeetree | |||||||||||||
| II. Leaves are once compound. | |||||||||||||
| A. Leaflets are trifoliate. | |||||||||||||
| PteleaHop Tree | |||||||||||||
| AA. Leaflets are always more than three. | |||||||||||||
| |. Leaflets are toothed. | |||||||||||||
| a. Stems are thorny. | |||||||||||||
| i. Plants have aromatic branches when crushed. | |||||||||||||
| ZanthoxylumPricklyash | |||||||||||||
| ii. Plants are without aromatic branches when crushed. | |||||||||||||
| GleditsiaHoneylocust | |||||||||||||
| aa. Stems are not thorny. | |||||||||||||
| i. Stipules are present, broad, and coarsely toothed. | |||||||||||||
| SorbusMountain-Ash | |||||||||||||
| ii. Stipules are absent. | |||||||||||||
| A) Pith is chambered. | |||||||||||||
| JuglansButternut, Walnut | |||||||||||||
| AA) Pith is solid. | |||||||||||||
| |) Buds are naked or valvate. | |||||||||||||
| a) Buds are naked. | |||||||||||||
| RhusSumac | |||||||||||||
| aa) Buds are valvate. | |||||||||||||
| CaryaHickory | |||||||||||||
| ||) Buds have several overlapping scales. | |||||||||||||
| CaryaHickory | |||||||||||||
| ||. Leaflets are entire. | |||||||||||||
| a. Stipules are absent or fall early. Side buds are visible. | |||||||||||||
| i. Buds are naked. | |||||||||||||
| RhusSumac | |||||||||||||
| ii. Buds are scaly. | |||||||||||||
| Ailanthus Tree of Heaven | |||||||||||||
| aa. Stipules are present. Side buds are usually hidden by petiole bases or sunken beneath leaf scars. | |||||||||||||
| RobiniaLocust | |||||||||||||
| Subgroup IILeaves are lobed. | |||||||||||||
| I. Lobes are entire and not regularly toothed. | |||||||||||||
| A. Buds are not clustered at the end of the twigs. | |||||||||||||
| |. Leaves on 2-year-old twigs are borne mostly in clusters of three to five on spurs. Leaves are fan-shaped with parallel veins. | |||||||||||||
| GinkgoGinkgo | |||||||||||||
| ||. Leaves on 2-year-old twigs are not borne in clusters on spurs. | |||||||||||||
| a. Stipular lines encircle the twig. | |||||||||||||
| LiriodendronTuliptree | |||||||||||||
| aa. Stipular lines are absent. | |||||||||||||
| i. Branches and foliage are fragrant when crushed. | |||||||||||||
| SassafrasSassafras | |||||||||||||
| ii. Branches and foliage are not fragrant when crushed. | |||||||||||||
| PopulusPoplar | |||||||||||||
| AA. Buds are clustered at the ends of the twigs. | |||||||||||||
| QuercusOak | |||||||||||||
| II. Lobes are toothed. | |||||||||||||
| A. Leaves are woolly or pubescent (hairy) beneath. | |||||||||||||
| PopulusPoplar | |||||||||||||
| AA. Leaves are not woolly or pubescent (hairy) beneath, except possibly on veins when young. | |||||||||||||
| |. Buds are hidden by the base of the petiole. Terminal bud is lacking. Stipule scar extends around the stem. | |||||||||||||
| PlatanusPlanetree | |||||||||||||
| ||. Buds are not hidden by petiole base. Stipule scars are not encircling the twig. | |||||||||||||
| a. Leaves are palmately veined and star-shaped. Older twigs usually have corky ridges. | |||||||||||||
| LiquidambarSweetgum | |||||||||||||
| aa. Leaves are pinnately veined and vary in shape from entire to mitten- shaped or lobed. Twigs are without corky ridges. | |||||||||||||
| MorusMulberry | |||||||||||||
| Subgroup IIILeaves are toothed. | |||||||||||||
| I. Buds are valvate. Leaves are mostly narrow. Veins are reticulate (curving and connected by cross veins). | |||||||||||||
| SalixWillow | |||||||||||||
| II. Buds have several overlapping scales. | |||||||||||||
| A. Twigs are thorny. | |||||||||||||
| CrataegusHawthorn | |||||||||||||
| AA. Twigs are not thorny, but may have stiff, short, spurs. | |||||||||||||
| |. Black glands (small, short, and threadlike) are present on midrib. | |||||||||||||
| MalusCrabapple | |||||||||||||
| ||. Black glands on the midrib are absent. | |||||||||||||
| a. Veins are straight, parallel, and running to the teeth without much branching. | |||||||||||||
| i. Outer bud scales number two to four, seldom five. | |||||||||||||
| A) Winter catkins are present. Spur-like branches are present on 2- to 3-year-old twigs, each bearing two to three leaves. Leaves are crowded almost in a whorl. | |||||||||||||
| BetulaBirch | |||||||||||||
| AA) Winter catkins and spur-like branches are absent. | |||||||||||||
| I) Leaves are two to three times as long as they are wide. Leaf margins are coarsely toothed and leaf tips are bristle-like. | |||||||||||||
| CastaneaChestnut | |||||||||||||
| II) Leaves are nearly as wide as they are long. Leaf bases are usually heart-shaped or lopsided. | |||||||||||||
| Tilia Linden | |||||||||||||
| ii. Outer bud scales number five or more. | |||||||||||||
| A) Buds are clustered at the ends of the terminal twigs. | |||||||||||||
| QuercusOak | |||||||||||||
| AA) Buds are not clustered at the ends of the terminal twigs. | |||||||||||||
| |) Buds are long-pointed with many scales. Leaves are either coarsely and singly toothed or simple with a wavy margin. | |||||||||||||
| FagusBeech | |||||||||||||
| ||) Buds are smaller and more ovate with fewer scales. Leaves are finely and doubly toothed. | |||||||||||||
| a) Leaves are rounded or heart-shaped at the base and rarely lopsided. | |||||||||||||
| I] Bud scales are marked with fine, longitudinal lines. Leaves have each side vein forming one or two branches near the margin of the leaf. | |||||||||||||
| OstryaHophornbeam | |||||||||||||
| II] Bud scales are without fine, longitudinal lines. Side veins are unbranched and run straight to the teeth. | |||||||||||||
| CarpinusHornbeam | |||||||||||||
| aa) Leaves are usually lopsided at the base. | |||||||||||||
| UlmusElm | |||||||||||||
| aa. Veins are anastomosing, branching, and curving before reaching the margin of the leaf. (reticulate) | |||||||||||||
| i. Leaf stalk is flattened vertically. | |||||||||||||
| PopulusPoplar | |||||||||||||
| ii. Leaf stalk is not flattened vertically but is round or nearly so. | |||||||||||||
| A) Terminal bud is sticky and usually much larger than the side buds. | |||||||||||||
| PopulusPoplar | |||||||||||||
| AA) Terminal or pseudo-terminal bud is not sticky and usually not larger than the side buds. | |||||||||||||
| |) Twigs have a bitter, almond taste. | |||||||||||||
| PrunusPlum, Cherry | |||||||||||||
| ||) Twigs do not have a bitter, almond taste. | |||||||||||||
| a) Terminal buds are present. | |||||||||||||
| I] Pith is diaphragmed or intermittently chambered. Leaves are prominently three-veined. Teeth are missing on lower third of the margin. Leaves are usually lopsided at the base. | |||||||||||||
| CeltisHackberry | |||||||||||||
| II] Pith is solid. Leaves are distinctly pinnately veined. Teeth are continued from base to tip or nearly so. Buds are long and pointed with four or five scales. | |||||||||||||
| AmelanchierServiceberry | |||||||||||||
| aa) Terminal bud is absent. | |||||||||||||
| I] Sap of freshly cut twigs is milky. Leaves are sometimes lobed. | |||||||||||||
| MorusMulberry | |||||||||||||
| II] Sap of freshly cut twigs is not milky. Leaves are finely toothed or nearly entire. | |||||||||||||
| OxydendrumSourwood | |||||||||||||
| Subgroup IVLeaves are entire. | |||||||||||||
| I. Leaves are narrow and evergreen-like. | |||||||||||||
| A. Leaves are spirally arranged or clustered on short, lateral spurs. | |||||||||||||
| LarixLarch | |||||||||||||
| AA. Leaves are alternate, spreading radially, or in two-ranks. Leaves are not borne on spurs. | |||||||||||||
| TaxodiumBald-Cypress | |||||||||||||
| II. Leaves are broader and not evergreen-like. | |||||||||||||
| A. Stems are thorny. Sap of freshly cut twigs is milky. | |||||||||||||
| MacluraOsage-Orange | |||||||||||||
| AA. Stems are not thorny. | |||||||||||||
| |. Stipules or their scars completely encircle the twig. | |||||||||||||
| MagnoliaMagnolia | |||||||||||||
| ||. Stipules or their scars do not encircle the twig. | |||||||||||||
| a. Twigs are aromatic when crushed or broken. | |||||||||||||
| i. Leaves are variable in shape. Leaves are lobed, mitten-shaped, or entire. | |||||||||||||
| SassafrasSassafras | |||||||||||||
| ii. Leaves are always entire. | |||||||||||||
| CotinusSmoketree | |||||||||||||
| aa. Twigs are not aromatic when crushed or broken. | |||||||||||||
| i. Buds are clustered at the ends of the terminal twigs. | |||||||||||||
| QuercusOak | |||||||||||||
| ii. Buds are not clustered at the ends of the terminal twigs. | |||||||||||||
| A) Branching is sympodial (side bud takes the lead). Leaves have parallel veins. | |||||||||||||
| CornusDogwood | |||||||||||||
| AA) Branching is normal or monopodial (end bud taking the lead). | |||||||||||||
| |) Leaves are palmately veined. Leaf shape is roundish or broadly ovate with heart-shaped leaf bases. | |||||||||||||
| CercisRedbud | |||||||||||||
| ||) Leaves are pinnately or indistinctly veined. | |||||||||||||
| a) Pith is diaphragmed or chambered. | |||||||||||||
| I] Terminal bud is present. | |||||||||||||
| B. Leaves are 243/4 inches (512 cm) long, glossy above, and somewhat bloomy beneath. | |||||||||||||
| NyssaTupelo | |||||||||||||
| BB. Leaves are 612 inches (15301/2 cm) long. Buds have only two to three scales and are woolly brown. | |||||||||||||
| AsiminaPawpaw | |||||||||||||
| II] Terminal bud is absent. | |||||||||||||
| DiospyrosPersimmon | |||||||||||||
| aa) Pith is solid. Leaves are usually finely toothed. | |||||||||||||
| OxydendrumSourwood | |||||||||||||